Quote ="Richie"I'm not sure you can "tone" a muscle. You can make a muscle stronger or weaker. If it gets stronger, it tends to get bigger (and certain types of strength have more affect on size than others) and if it gets weaker it gets smaller. Then you can gain or lose fat. '"
I am not claiming to be an expert in this but I don't think that is basically right. Resistance training - eg weights- leads to hypertrophy -i.e. a large increase in the number of myofilaments - and this is what increases the visible size and shape of muscles. The basic reason is that they are being asked to do more of one particular type of work (the resistance in that particular direction) and so repetitions lead the body to do that, so the muscle can be stronger, so it can do the work more efficiently.
However for long distance running what the muscle needs is endurance rather than strength. I am of course certain that Mo Farah's muscles are far stronger than the majority of the population, and certainly plenty strong enough to propel him forward at his pace, but they don't need to be any bigger, and if they were, they'd weigh more, so he would suffer a weight-related handicap. (Possibly his glutes excepted as they provide most of the driving force). They need to be strong enough to absorb the repeated landing and bounce off the running surface for the next stride.
Quote ="Richie"You're certainly right about gaining muscle mass, and beyond a certain point you can't do it without being on a calorific surplus. Which is how I've come to learn about gaining and losing weight lately after too much time trying to gain strength and lose fat at the same time, to little effect.'"
I have no link for this, but I'm pretty sure that long distance runners usual routine in the "off season" does include much more strength work to increase strength and muscle size/mass, but they then aim to reduce that mass/weight of muscle by the start of the season back to their fighting weight.